Recent Studies


Multi-level control of adaptive camouflage by European cuttlefishCurrent Biology (with  D. Osorio, F. Ménagerm and A. Darmaillacq). PDF Additional Commentary

The nature of illusions: A new synthesis based on verifiability. Frontiers in Human Neuroscience, 16, 2022. PDF

Brain trauma impacts retinal processing: photoreceptor pathway interactions in traumatic light sensitivity. Documenta Ophthalmologica, 144(2):1-12, 2022 (with L.T. Likova). PDF

Guest editorial: On the power and legitimacy of follow-up testing. Perception 51:225–229. PDF

Self-regulation of seat of attention into various attentional stances facilitates access to cognitive and emotional resources: An EEG study. Frontiers in Psychology, 24 February 2022 (with G Hartelius and LT Likova). PDF


A gain-control disparity energy model for perceived depth from disparity. Vision Research, 181, 38–46, 2021 (with PY Chen and CC Chen). PDF

Cartography as spatial representation: A new assessment of the competing advantages and drawbacks across fields of science. Journal of Perceptual Imaging, 2021:1561-15610. PDF

A live experience of four-dimensional structure. Perception, 50(2): 165-169, 2021. PDF

Did Tim paint a Vermeer? Electronic Imaging, 2021, 60403–1, 2021 (with DG Stork and SJ Schechner). PDF

The interstitial pathways as the substrate of consciousness: A new synthesis. Entropy, 23, 1443, 2021. PDF


The science of Renaissance perspective. Perception, 49(12), 2020. PDF

Ten testable properties of consciousness. Frontiers in Psychology, 11:1144, 2020.  PDF

Are there four, six, or seven unique hues? VSS 2020. Poster Video

An accelerated cue combination principle accounts for multi-cue depth perception.
Journal of Perceptual Imaging
. 3(1): 010501-1–010501-9, 2020. PDF

The fundamental nature of time. Journal for Research in Philosophy and History, 3:(1), 2020. PDF


Dynamic amodal completion through the Magic Wand Illusion. i-Perception. 10(6):2041669519895028. PDF

Measurement of saccadic eye movements by electrooculography for simultaneous EEG recording. Behav Res Methods. 51:2139-2151. 2019  (with Y Jia). PDF

Length summation in noise. Journal of Vision, 19(9):11, 1–13, 2019 (with CC Chen and YH Yeh). PDF

Eye centring in selfies posted on Instagram. PLoS ONE 14(7): e0218663 (with N Bruno and M Bertamini). PDF (also see interview here and articles)

Guest editorial: Depth cue combination: A quantitative critique. Perception 48:765–768, 2019. PDF

Evidence that Leonardo da Vinci had strabismus. JAMA Ophthalmol. Jan 1;137(1):82-86, 2019.  PDF (also see links to article coverage.)

Points of contact between the Stappian philosophy and emergent aspect dualism. Activitas Nervosa Superior, 81:1-6, 2019.  PDF


The emergent aspect dualism view of quantum physics: a new ontology to resolve the complementarity conundrum. J Res Phil Hist 1:166–182, 2018. PDF 

Development and validation of a new glaucoma screening test using temporally modulated flicker. Ophthalmic Physiol Opt. 38:617-628, 2018. (with Fidalgo BR, Jindal A, Ctori I, Lawrenson JG.) PDF

A Brücke-Bartley effect for contrast. Roy Soc Open Sci. 5:180171, 2018. eCollection 2018. (with Solomon JA) PDF

Rational approaches to correcting for multiple tests. Human Vision and Electronic Imaging (535) 1-8, 2018. PDF

Absolute distance from perspective. Perception 47:581-584, 2018. PDF

When light hurts: Comparative morphometry of human brainstem in traumatic photalgia. Scientific Reports, 8(1), 2018 (with LT Likova). PDF.

Does colour filling-in account for colour perception in natural images? i-Perception, 9(3), 2018 (with JA Solomon). PDF.


Computational estimation of scene structure through texture gradient cues. Human Vision and Electronic Imaging  (10)167-176, 2017 (with A Gopi). PDF

Studying the retinal source of photophobia by facial electroretinography. Optom Vision Sci 94:511-518 , 2017 (with LT Likova). PDF

Improvement of contrast sensitivity with practice is not compatible with a sensory threshold account. J Opt Soc Amer 34:870-880, 2017 (with JA Solomon). PDF


Peripheral color vision and motion processing. Human Vision and Electronic Imaging, 5:1-5, 2016. PDF

The cortical network for Braille writing in the blind. Human Vision and Electronic Imaging  6:1-6, 2016 (with LT Likova, L Cacciamani, K Mineff, SC Nicholas). PDF

The perceived depth from disparity as function of luminance contrast. Journal of Vision, 16(11):20, 1–9, 2016 (with P-Y Chen, C-C Chen). PDF

Editorial: Neural signal estimation in the human brain. Front Neurosci 10:185 eCollection 2016. (with C Howarth, LT Likova). PDF

Evidence of stereoscopic surface disambiguation in the responses of V1 neurons. Cereb Cortex 27:2260-2275, 2016 (with JM Samonds, TS Lee). PDF

Inefficiency of orientation averaging: Evidence for hybrid serial/parallel temporal integration. J Vision 16:13, 2016 (with JA Solomon, KA May). PDF


Deficits in the activation of human oculomotor nuclei in chronic traumatic brain injury. Front Neurol 6:173, 2015 (with LT Likova, KN Mineff, SC Nicholas SC). PDF

Shading beats binocular disparity in depth from luminance gradients: Evidence against a maximum likelihood principle for cue combination. PLoS One 10:e0132658, 2015 (with CC Chen). PDF

Analysis of neural-BOLD coupling through four models of the neural metabolic demand. Front Neurosci. 9:419, 2015 (with LT Likova and SC Nicholas). PDF

Short and sweet: peripheral color demo. i-Perception 6: 2041669515613671, 2015. PDF

Consequences of traumatic brain injury for human vergence dynamics. Front Neurol 5:282, 2015 (with LT Likova, KN Mineff, AM Elsaid, SC Nicholas). PDF

The vault of perception: Are straight lines seen as curved? Art & Perception 3:117-137, 2015. PDF

Effect of familiarity on Braille writing and reading in the blind: From graphemes to comprehension. J Vision 15:920-920, 2015 (with LT Likova, L Cacciamani, KN Mineff, SC Nicholas). PDF 


Review: Possibilities of perception. Perception 43, 353-354, 2014. PDF

The neurometabolic underpinnings of fMRI BOLD dynamics. In Advanced Brain Neuroimaging Topics in Health and Disease – Methods and Applications. Editor Papageorgiou TD, Christopoulos GI, Smirnakis SM. Intech Press, Ch 2, 2014, 21-47, 2014 (with LT Likova). PDF

Visual function, traumatic brain injury, and posttraumatic stress disorder. Journal of Rehabilitation Research & Development 51 (4), 547, 2014 (with GL Goodrich, GL Martinsen, HM Flyg, J Kirby, DW Garvert). PDF

The autostereogram, Scholarpedia, 2014. PDF

Space perception as embodied cognition. Human Vision and Electronic Imaging XIX, 2014. PDF

Bela Julesz: A memoir. Proc Nat Acad Sci. 110, 2014. PDF

Is the mind in the head? A belated response to Koenderink (1999). Perception guest editorial 43 (2), 103-106, 2014. PDF

Leonardo da Vinci’s world map. In Huylebrouck D (Ed), Leonardo’s Mathematics, Springer: Berlin, 2014. PDF


Binocular eye movements in health and disease. Human Vision and Electronic Imaging XVII, 8651-34, 2013. PDF

Dips and Bumps: On Bloch’s law and the Broca-Sulzer phenomenon. Proc Nat Acad Sci. 110:E1330, 2013 (with A Gorea). PDF

Depth structure from asymmetric shading supports face discrimination. PLoS One 8(2): e55865, 2013 (with Chen C-C, Chen C-M). PDF

Recurrent connectivity can account for the dynamics of disparity processing in V1. J Neurosci 33 2934-2946, 2013 (with JM Samonds, BR, Potetz, TS Lee). PDF

A simpler structure for local spatial channels revealed by sustained perifoveal stimuli. J Vision 13:1, 22, 2013 (with LL Kontsevich). PDF


Brain mapping: The (un)folding of striate cortex. Current Biology 22(24), R1051 – R1053, 2012 (with MM Schira, MGP Rosa). PDF

Analysis of human vergence dynamics. J Vision 12, 21, 2012 (with AM Elsaid, LT Likova, N Gill, SC Nicholas). PDF

Collinear facilitation over space and depth. J Vision 12. pii: 20, 2012 (with PC Huang, CC Chen). PDF

3D discomfort from vertical and torsional disparities in natural images. Human Vision and Electronic Imaging XVII, 2012 (with LT Likova, K Atanassov, V Ramachandra, S Goma). PDF

The role of the visual arts in enhancing the learning process. Frontiers in Human Neuroscience 6:8 2012 (with LT Likova). PDF


Estimating neural signal dynamics in the human brain. Frontiers in Systems Neuroscience 5:33, 2011 (with LT Likova). PDF

Perceptual coding for 3D reconstruction. EUVIP 2011 10.1109/EuVIP.2011.604553, 2011 (with SC Nicholas). PDF

The role of disparity interactions in the perception of the 3D environment. In Vision in 3D Environments, LR Harris and MRM Jenkins, Eds. Cambridge University Press, p 95-114, 2011. PDF

Paradoxical perception of surfaces in the Shepard tabletop illusion. i-Perception 2, 137–141, 2011. PDF

The role of midlevel surface representation in 3D object encoding. In Computer Vision: From Surfaces to 3D Objects. Tyler CW, (Ed) Chapman Hall: New York, p vii-xxiii, 2011. PDF

Visual surface encoding: A neuroanalytic approach. In Computer Vision: From Surfaces to 3D Objects. Tyler CW (Ed) Chapman Hall: New York, p 163-182, 2011 (with LT Likova). PDF


Modeling magnification and anisotropy in the primate foveal confluence. PLoS, Computational Biology 6(1):e1000651, 2010 (with MM Schira, M Breakspear, B Spehar). PDF

Symmetry: modeling the effects of masking noise, axial cueing and salience. PLoS One 5(4):e9840, 2010 (with CC Chen). PDF

An algebra for the analysis of object encoding. NeuroImage, 50:1243-50, 2010 (with LT Likova). PDF

Metameric intransitivity. Att Percept Psychophys, 72:891-93, 2010 (with AE Huang, AJ Hon, EL Altschuler).  PDF